Ever-increasing interest in oceanography and marine biology and their relevance to global environmental issues create a demand for authoritative reviews summarizing the results of recent research. Oceanography and Marine Biology: An Annual Review has catered to this demand since its founding by the late Harold Barnes more than 50 years ago. Its objectives are to consider, annually, the basic areas of marine research, returning to them when appropriate in future volumes; to deal with subjects of special and topical importance; and to add new subjects as they arise. The favourable reception and complimentary reviews accorded to all the volumes shows that the series is fulfilling a very real need. Volume 55 follows closely the objectives and style of the earlier volumes, continuing to regard the marine sciences-with all their various aspects-as a unity. Physical, chemical, and biological aspects of marine science are dealt with by experts actively engaged in these fields, and every chapter is peer-reviewed by other experts working actively in the specific areas of interest. Chapters 3, 4, 5, and 6 of this book are freely available as downloadable Open Access PDFs at http://www.taylorfrancis.com under a Creative Commons Attribution-Non Commercial-No Derivatives (CC-BY-NC-ND) 4.0 license.